The Botany of Iceland - 01.12.1912, Page 159

The Botany of Iceland - 01.12.1912, Page 159
MARINE ALGAL VEGETATION 145 mediate forms, so are the various associations connected. If the deep-water character of the association is taken as a starting point, we notice that it changes gradually with decreasing depths on ex- posed coasts, and tliat somewhere near the limit of low-tide it assumes an entirely different characler, which is displayed in tlie leathery, long-stemmed form with a narrow and slightlj'-divided frond (f. stenophi]lla). On an exposed coast this variety might be termed the shallow-water type or perhaps, rather, tlie surf-type, in conforniity with the surf-form of Fucus inflatus. The surf-character was very beautifully and typically developed in the Vestmannaeyjar; here f. stenophytla grew very socially, forming a continuous belt, the upper boundary of which occurred almost at the limit of low- tide. When during low-tide the waves receded it was very inter- esting to see how easily the leathery, narrow, slightlv-divided laminæ moved with the waves, and everywhere, as far as the waves receded, tlie rocky substratum between the StenophyZ/a-individuals was quite reddish in colour from the encrusting Phymatolithon polymorphum. The Laminaria plants were very firmly attaclied to the rock — a fact evidently well known to the fishermen, as tliey fastened the boat to a Laminaria while we went ashore. Tlie stipe is leathery and pliable, and the plants cling closely to the substratum when the waves recede. I liave found /'. stenophylla in other places, although not in such abundance, and not quite so typical. Here the same rule applies as with regard to Fucus inflatus and Laminaria saccharina, that the tendency to vary seems to depend on the greater or lesser movement of the water; on coasts which are somewhat exposed, a Laminaria digitata-association is rather frequently found at about the limit of low-tide, with a character midway hetween the surf- character and the deep-water character. If we again take the deep-water character of the association as a starting point, and move inwards towards the protected coasts, we see that the character changes again, but in another direction. The slipe becomes shorter and the frond much broader and slit into fewer and broader lobes. Within the fjords, in W. and E. Ice- land especially, the character is entirely different from the deep- water character. Here occur forms with very broad fronds which are either undivided or divided into two, or a few, very broad lobes (f. cucullata). Generally the depth is about 4—20 metres, even deeper occasionally. I have found associations with this character The Botany of Iceland. I. 10
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