Fróðskaparrit - 01.07.2004, Síða 54

52
WETLAND COLEOPTERA OF THE FAROE ISLANDS
ence to work done in 2002; here it is suffi-
cient to note that this difference is less
clear, if anything the reverse, in northern
Scotland. The widespread occurrence of
Agabus bipustulatus, with larvae present in
all still and most slow-running habitats, is
not typical of areas occupied by other large
diving beetles.
The prediction of lentic species in the
Faroes is within 0.0068 logarithmic units of
the actual 12 species (Sphaeridiinae ex-
cluded). This is the lowest residual of all
eighteen areas under consideration, possi-
bly reflecting the accuracy of the observa-
tion for the Faroes.
Finally, there is no evidence of beetles
responding to climate change. In Scotland,
the most extreme example thus far is that of
Rhantus suturalis (Macleay). This species,
which ranges from Europe to New Zealand,
was known until recently in Scotland only
from the extreme south on six occasions up
to 1950. It was discovered in Fife by Mr T.
Huxley in 2002, and in 2003 by GNF near
the northern extremity of mainland Scot-
land in Caithness, at a site that had been
sampled frequently since 1985. Thus there
is activity further south that may yet result
in additions to the Faroese fauna.
Acknowledgements
We are grateful to Dr Ignacio Ribera for re-
working the analysis of the species-area re-
lationship. GNF wishes to note with grati-
tude the provision of facilities for this
work. Rob Close accompanied us on the
survey work in June 2004. Derek Lott iden-
tified the Staphylinidae.
References
Angus, R.B. 1970. A revision of the beetles of the genus
Helophorus F. (Coleoptera, Hydrophilidae) subgen-
era Orphelophorus d’Orchymont, Gephelophorus
Sharp and Meghelophorus Kuwert. Acta zoologica
Fennica 129: 3-62.
Angus, R.B. 1982. Separation of two species standing as
Helophorus aquaticus (L.) (Coleoptera, Hydrophili-
dae) by banded chromosome analysis. Systematic
Entomology 7: 265-281.
Angus, R.B. 1985. Towards a revision of the Palaearctic
species of Helophorus F. (Coleoptera, Hydrophili-
dae). 1. Entomologicheskoe Obozrenie 64: 716-747
(In Russian-English version as Entomological Re-
view (1986) 64 (4): 128-162).
Angus, R.B. 1992. Insecta: Coleoptera: Hydrophilidae:
Helophorinae. Susswafierfauna von Mitteleuropa 20
(10): part 2, 144 pp. Gustav Fischer Verlag.
Bengtson, S-A. 1981. Terrestrial invertebrates of the
Faroe Islands: III Beetles (Coleoptera): check-list,
distribution, and habitats. Fauna Norvegica Series B
28: 52-82.
Berge Henegouwen, A. van, 1986. Revision of the Eu-
ropean species of Anacaena Thomson (Coleoptera:
Hydrophilidae). Entomologica scandinavica 17:
393-407.
Brown, E.S. 1945. The Corixidae of the Faeroe Islands,
with observations on the geographical distribution
of Corixidae in neighbouring parts of the World.
Proceedings of the Zoological Society of London
114 (4): 490-506.
Drotz, M.K., Saura, A. and Nilsson, A.N. 2001. The
species delimitation problem applied to the Agabus
bipustulatus complex (Coleoptera, Dytiscidae) in
north Scandinavia. Biological Journal of the Lin-
nean Society 73: 11-122.
Falkenstrom, G. 1922. Stockholmstraktens Vatten-
Coleoptera. Entomologisk Tidskrift 43: 10-32.
Hansen, M. 1987. The Hydrophiloidea (Coleoptera) of
Fennoscandia and Denmark. Fauna entomologica
scandinavica 18, 254 pp., Leiden and Copenhagen.
Hansen, M. 1999. World Catalogue of lnsects Volume2.
Hydrophiloidea (s. str.) (Coleoptera). Apollo
Books, Stenstrup.
Hansen, M.J. 1881. Faunula Insectorum Faeroeensis.
Naur Tidsskift 13 (3): 229-250.
Holdhaus, K. 1911. Zur Kenntnis der Coleopteren-Fau-
na der Faroer. Deutsche entomologische National-
bibliotek, Berlin 2: 123-125.
Jackson, D.J. 1956. The capacity for flight of certain
water beetles and its bearing on their origin in the