Fróðskaparrit - 01.01.2008, Page 180

Fróðskaparrit - 01.01.2008, Page 180
178 DISTRIBUTION AND THE IMPACT OF OUTFIELD DRAINAGE ON CARABIDS (COLEOPTERA, CARABIDAE) IN NORTH WESTERN EYSTUROY, FAROE ISLANDS tudes in the ND area, N. rufescens domina- ting the more stony areas while N. salina is dominating areas with less gravel and stones and more grass, which is in good accordance with results from Danielsen and Hansen (2000), Sadler and Dugmore (1995) and de- scribtions by Lindroth (1986). The reason for Nebria salina being sig- nificantly more abundant at the higher alti- tudes in UD area - even though the lower areas are suitable - might be competition from Calatbus fuscipes at the lower altitudes. These two species are quite similar in size Ca- iathus fuscipes being a bit larger (10-14.4 mm) than Nebria salina (10-13.5 mm) (Lin- droth, 1985; 1986). General principles of co- existence predict that species that are equal in size do not share the same habitat if they use the same resourses. Bengtson (1982) working with habitat utilisation and niche breadths found that there was very little overlap between Nebria salina and Calathus fuscipes. This, together with our results point to C.fuscipes excluding N. salina from areas where C.fuscipes dominates. An explanation for Nebria rufescens being more abundant at the higer altitudes in the UD area can be re- lated to the riversides and banks having more stones and gravel in the higher altitude areas, this is one of Nebria rufescens' pref- ered habitats (Lindroth, 1985) and also cor- responds well with results from Danielsen and Hansen (2000). The upper altitude areas in the OD area contain more heather, while grasses become increasingly more dominant in the lower al- titudes, which could make the lover altitudes better suitable to Nebria salina and Nebria rufescens, compared to the upper altitudes (Lindroth, 1985). The OD area shows sign of drying up (Fosaa et al., 2008); therefore dif- ferences in distribution according to alti- tudes that might have been are cancelled be- cause the upper heather influenced areas have become drier (Fosaa et al., 2008), which makes them better suited to Nebria salina and Nebria rufescens (Lindroth, 1985). Also one of the rivers that pit-fall traps were placed along in this area did not have so much heather at the upper altitudes com- pared to the lower altitudes, which mini- mizes differences in habitat structure. Hansen (2006) also found more Trechus obtusus at higher altitudes; the present find- ing of higher abundance of T. obtusus at the higher altitudes in the UD area might be re- lated to the fact that the lower elevations in the UD area are a lot steeper than the higher altitudes, therefore the lower elevation might be drier as a whole. Also the lower al- titudes in UD area appeared to be heavily grazed because of the very short vegetation and high presence of sheepdroppings; and therefore minimizing shelter in the vegeta- tion. All this reduces the favourable condi- tions to Trechus obtusus at the lower alti- tudes (Lindroth, 1985). The association between vegetation and carabidae in the 3 areas is summarized in Table 5. Typically Nebria salina, Nebria rufescens, Patrobus septentrionis, Patrobus atrorufus and Trechus obtusus were among the domi- nating species. Carabus problematicus was only among the dominating in the OD area and in the lower altitudes in the UD area and Calathus fuscipes dominated in the lower al- titudes in the OD and UD area (Tables 2, 3 and 4). The higher abundance of Carabus problematicus in the OD area is related to
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