Náttúrufræðingurinn - 2005, Page 45

Tímarit Hins íslenska náttúrufræðifélags
forefather in Ukrania as previously sug-
gested. The questions of when and how
the horse was domesticated and tamed
are discussed and compared to the histo-
ry of domestication of the other farm ani-
mals. A chapter on grazing and the
nature of horse feeding and digestion
compared to ruminants discusses the
relationship between foraging behav-
iour, anatomy and digestive system in
evolutionary terms. Finally, social behav-
iour of the species is discussed empha-
sizing how the environment has shaped
their behaviour throughout their evolu-
tionary history. Also, the two social sys-
tems which characterize different species
within the genus, ie Type I (dominant
male with a harem of females and sub-
adults) and Type II (with males defend-
ing mating territories) are discussed in
light of selection factors and lastly how
the behaviour of the domestic horse has
remained remarkedly unchanged.
Heimildir
1. MacFadden, B.j. 1992. Fossil horses. Systematics, paleobiology, and
evolution of the family Equidae. Cambridge University Press. 369 bls.
2. Forstén, A. 1992. Mitochondrial-DNA time table and the evolution of
Equus: comparison of molecular and paleontological evidence. Ann.
Zool. Fennici 28. 301-309.
3. Oakenfull, E.A., Lim, H.N. & Ryder, O.A. 2000. A survey of equid
mitochondrial DNA: Implications for the evolution, genetic diversity
and conservation of Equus. Conserv. Genetics 1. 341-355
4. Orlando, L., Eisenmann, V., Reynier, F., Sondaar, P. & Hanni C. 2003.
Morphological convergence in Hippidion and Equus (Amerhippus) South
American Equids elucidated by ancient DNA analysis. J. Molecular
Evolution 57. S29-S40.
5. MacFadden, B.J. & Shockey B.J. 1997. Ancient feeding ecology and
niche differentiation of Pleistocene mammalian herbivores from Tarija,
Bolivia: Morphological and isotopic evidence. Paleobiology 23(1).
77-100.
6. Martin, P.S. & Wright, H.E., Jr. (ritstj.). 1967. Pleistocene Extinctions:
The search for a cause. New Haven. Yale University Press.
7. Martin, P.S. & Klein, R.G. (ritstj.). 1984. Quarternary Extinctions: A
Prehistoric Revolution. Tucson. University of Arizona Press.
8. Azzaroli, A. 1991. Ascent and Decline of monodactyl Equids - a case for
prehistoric overkill. Annales Zoologici Fennici 28(3-4). 151-163.
9. Alberdi, M.T., 2001. Hippidioti saldiasi Roth 1899 (Equidae, Perisso-
dactyla), at the Piedra Mueo Site (Santa Cruz, Argentina): Its
implication for the regional economy and environmental recon-
struction. J. Archaeological Sci. 28. 411-419.
10. Jansen, T., Forster, P., Levine, M.A., Oelke, H., Hurles, M., Renfrew, C.,
Weber, J. & Olek, K. 2002. Mitochondrial DNA and the origins of the
domestic horse. PNAS 99. 10905-10910.
11. Oakenfull, E.A., Lim, H.N. & Ryder, O.A. 2000. A survey of equid
mitochondrial DNA: Implications for the evolution, genetic diversity
and conservation of Equus. Conserv. Genetics 1. 341-355.
12. Vilá, C., Leonard, J.A., Götherström, A., Marklund, S., Sandberg, K.,
Lindén, K., Wayne, R.K. & Ellegren, H. 2001. Widespread origins of
domestic horse lineages. Science 291. 474-477.
13. Budiansky S. 1998. The nature of horses. Their evolution, intelligence
and behaviour. Phoenix - Orion Books Ltd. London.
14. MacHugh, D.E. & Bradley, D.G. 2001. Livestock genetic origins: Goats
buck the trend. PNAS 98. 5382-5384.
15. Vera, F.W.M. 2000. Grazing ecology and forest history. CABI
Publishing. 506 bls.
16. Van Dierendonck, M.C. & Wallis de Vries, M.F. 1996. Ungulate rein-
troductions: Experiences with the takhi or Przewalski horse (Equus
ferus Przewalski) in Mongolia. Conserv. Biol. 10. 728-740.
17. Feh, C. 2002. Relationships and communication in socially natural
horse herds. Dorothy Russell Havemeyer Foundation Workshop Horse
Behavior and Welfare 13-16 June 2002.
(http: / / www2.vet.upenn.edu / labs / equinebehavior / hvnwkshp / hv02 / hv02prog.
htm)
18. L. Boyd & N. Bandi 2002. Reintroduction of takhi, Equus ferus
przewalskii, to Hustai National park, Mongolia: time budget and
synchrony of activity pre- and post-release. Appl. Anim. Behav. Sci. 78.
87-102
19. King, S.R.B. 2002. Home range and habitat use of free-ranging Prze-
walski horses at Hustai National Park, Mongolia. Appl. Anim. Behav.
Sci. 78. 103-113.
20. (http://www.imh.org/imh/bw/zebra.html) (20.10.2003)
21. Feh, C., B. Munkhtuya., S. Enkhbold & T. Sukhbaatar 2001. Ecology and
social structure of the Gobi khulan Equus hemionus subsp. in the Gobi B
National Park, Mongolia. Biol. Conservation 101. 51-61.
22. Pough, F.H., Janis, C.M. & Heiser, J.B. 2005. Vertebrate Life. 7. útg.
Pearson Prentice Hall. New Jersey.
23. Janis, C. 1976. The evolutionary strategy of the equidae and the origins
of rumen and cecal digestion. Evolution 30. 757-774.
24. MacFadden, B.J. & Ceding, T.E. 1994. Fossil horses, carbon isotopes and
global change. Trends Ecol. Evol. 9. 481—486.
25. Goodwin, D. 2002. Horse behaviour: Evolution, domesticaton and feral-
isation. í: The Welfare of Horses (ritstj. N. Waran). Kluwer Acad. Pub.
26. Janis, C.M., Damuth J. & Theodor J.M. 2002. The origins and evolution
of the North American grassland biome: the story from the hoofed
mammals. PALAEO 177.183-198
27. Huston, J.E. & Pinchak, W.E. 1991. Range animal nutrition. Bls. 27-63 í:
Grazing management. An ecological perspective. (ritstj. R. K.
Heitschmidt & J.W. Stuth). Timber Press. Portland, Oregon.
28. McDonald, P., Edwards, R.A., Greenhalgh, J.F.D. & Morgan, C.A. 1995.
Animal nutrition (5. útgáfa). Longman Sci & Tech., UK.
29. Church, D.C. 1988 (ritstj.). The ruminant animal. Digestive physiology
and nutrition. Prentice Hall, New Jersey.
30. Cunha, T. J. 1991. Horse feeding and nutrition (2. útg.). Acad. Press, Inc.
31. Van Soest, P. J. 1982. Nurtitional ecology of the ruminant. O & B Books,
Oregon.
32. Hoffmann, R.R. 1988. Anatomy of the gastro-intestinal tract. Bls. 14-44
í: The ruminant animal. Digestive physiology and nutrition. (ritstj. D.C.
Church). Prentice Hall, New Jersey.
33. McMahon, T.A. & Bonner, J.T. 1983. On size and life. New York:
Scientific American Library.
34. Kleiber, M. 1975. The fire of life. R.E. Krieger Publ. Co. N. Y.
35. Thorhallsdottir, A.G. & Thorsteinsson, I. 1993. Behaviour and plant
selection. Icel. Agr. Sci. 7. 59-77.
36. Frape, D. 1986. Equine nutrition and feeding. Longman Sci & Tech.,
UK.
37. Cymbaluk, N.F. 1990. Comparison of forage digestion by cattle and
horses. Can. J. Anim. Sci. 70. 601-610.
38. Anna Guðrún Þórhallsdóttir & Tryggvi Eiríksson 1996. Samanburður á
meltanleika gróffóðurs hjá hrossum og sauðum. Rit Ráðunautafundar.
Bls. 189-193.
39. Anna Gudrun Thorhallsdottir, Gudni Agustsson & Johann Magnusson
2002. Grazing Behaviour of the Icelandic Horse. Dorothy Russell
Havemeyer Foundation Workshop Horse Behavior and Welfare 13-16
June 2002.
(http: / / www2.vet.upenn.edu/labs/equinebehavior/hvnwkshp/hv02/hv02prog.
htm)
40. Duncan, P., Foose, T.J., Gordon, I.J., Gakahu, C.G. & Lloyd, M. 1990.
Comparative nutrient extraction from forages by grazing Bovids and
Equids. A test of the nutritional model of Equid Bovid competition and
coexistence. Oecologia 84. 411-418.
41. Hintz, H.F. & Cymbaluk, N.F. 1994. Nutrition of the horse. Ann. Rev.
Nutr. 14. 243-267.
42. Gwynne, M.D. & Bell, R.H.V. 1968. Selection of vegetation components
by grazing ungulates in the Serengeti National Park. Nature 220.
390-393.
43. Levine, M.A. 2002. Domestication, breed diversification and early
history of the horse. Dorothy Russell Havemeyer Foundation
Workshop Horse Behavior and Welfare 13-16 June 2002.
(http: / / www2.vet.upenn.edu / labs / equinebeha vior / h vnwkshp / hv02 / hv02prog.
htm )
44. Brown, D.R. & Anthony, D.W. 1998. Bit wear, horseback riding, and the
Botai site in Kazakstan. J. Archaeol. Sci. 25. 331-347.
45. Anthony, D.W. & Brown, D.R. 1991. The origins of horseback riding.
Antiquity 246. 22-38.
46 Anthony, D.W. 1998. The opening of the Eurasian steppe at 2000 BCE.
í: The Bronze Age and Early Iron Age Peoples of Eastern Central Asia
(ritstj. V.H. Mair). Univ. Pennsylvania Pub. Phil.
47. Levine, M. 1999. Investigating the origins of horse domestication.
Equine Veterinary J. Suppl. 28. 6-14.
48. Mallory, J.P. 1989. In search of the Indo-Europeans. Thames and
Hudson, London.
49. Seder, M.A. & Hesse, B. 2000. The initial domestication of goats (Capra
hircus) in the Zagros mountains 10,000 years ago. Science 287.2254-2257.
50. Bruford, M.W., Bradley, D.G. & Luikart, G. 2003. DNA markers reveal
the complexity of livestock domestication. Nat. Rev. Gen. 4. 900-910.
51. Hiendleder, S., Kaupe, B., Wassmuth, R. & Janke A. 2002. Molecular
analysis of wild and domestic sheep questions current nomenclature
and provides evidence for domestication from two different
subspecies. Proc. Royal Soc. Lond. Series B-Biol. Sci. 269. 893-904.
52. Hermanson, J.W. & MacFadden, B.J. 1996. Evolutionary and functional
morphology of the knee in fossil and extant horses (Equidae). J. Vert.
Paleontol. 16. 349-357.
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