Fróðskaparrit - 01.01.2007, Side 186

Fróðskaparrit - 01.01.2007, Side 186
184 THE IMPACT OF GRAZING ON MOUNTAIN VEGETATION AND THE ARBUSCULAR MYCORRHIZAL SYMBIONT Discussion Several short term studies of exclusions of herbivores have shown that the vegetation biomass increases (e.g. Austrheim and Eriksson, 2001). We found some changes in the vegetation, some of them significant, as a result of sheep grazing. During the study period, the biomass increased con- siderably in the enclosures, which can be seen by the increase in length of grass- leaves and vegetation cover. Our short term results gave a significant increase in both cover and vegetation height in the lowland site as well as the alpine site for thc ungrazed plots. As the vegetation cover and the vegetation height is much higher in lowland plots than in the alpine plots it would be expected that the gain in biomass was higher in the lowland site compared to the alpine site. This was also the case, but the difference was much smaller than would be expected. We must also take into consideration here that the alpine site was sampled after five years of enclosure and the lowland site after six years of enclosure. These fmdings indicate that the alpine site also is a productive site (Korner, 1999) and that the herbivores have a profound effect on the alpine eco- system. We have seen that five plant species changed their abundance significantly. Only small changes in loss and gain of new species had occurred at this stage, and maybe the time to detect such changes must be longer. The abundances of graminoids increased in grazed plots in the lowland while the opposite was the case in the alpine area. In the lowland plots, this increase was due to increased abundance of the non-palatable Nardus stricta while other grass species were decreasing. This fínding could be explained by competition between species instead of the grazing ef- fect. This is also seen by others (e.g. Welch and Scott, 1995). At the alpine site, Nardus stricta was not found, but here all grass species increased their abundance in the enclosures. This result can be explained by the stronger effect of grazing in areas of low biomass and lesser effect due to com- petition (van der Wal et. al., 2000). In the Scottish heathland, a 20-years study showed that Calluna declined at four sites where there were increases in herbi- vores. At these sites, graminoids and herbs showed a general rise in cover, and eri- coids and lichens decreased. Species in- creasing significantly included Agrostis capillaris, Anthoxantum odoratum, Fes- tuca ovina, Galium saxatilis, Luzula multi- flora, Nardus stricta and the moss Rhytidi- adelphus squarrosus. In contrast, De- schampsia flexuosa was reduced in cover at the same sites (Welch and Scott, 1995). The response of Agrostis capillaris to grazing is variable. Some studies have found the abundance of Agrostis capillaris to decrease when grazed (Magnússon and Magnússon, 1990; Crawley, 1990). Ac- cording to Magnússon and Magnússon (1990), the abundance and cover of Agrostis capillaris decreases with increas- ing grazing intensity. This study from Ice- land might be the best predictor for what to expect in the Faroe Islands, as the climatic conditions are quite similar. Other studies have found the abundance
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