Skógræktarritið - 15.05.2001, Qupperneq 151

Skógræktarritið - 15.05.2001, Qupperneq 151
Table I. Examples of some of the important processes to be included in the HIBECO mountain birch forest ecosystem model. Suggestive links to their respec- tive characteristic scales (response times and spatial correlation lengths) are also given. The spatial and temporal aspects of a given factor may be correlated (not shown). Preliminary definitions of scale: Fine spatial scale, approx. 500-2000 m2; Coarse spatial scale, approx. 100-200 km and beyond; fine temporal scale: approx. I day; coarse temporal scale: approx. one season or longer. Birch growth Space: Fine scale processes: Local exposition, altitude, soil quality, local climate conditions Coarse scale nrocesse.s: Global warming effects, provenance differ- ences and regional adaptation (e.g., frost tolerance of buds) Time: Fine sc.ale processes: Episodic influence from frost and herbivory Coarse scale processes: Dieback due to repeated defoliation from insect attack over consecutive years (threshold effect); herbivory inhibitory response (delayed density dependence) on expense of maximum growth rate, successional stage effects. Herbivory Space: Fine scale nroc.esses: Sheep as a fine-scale forager (stand edges vs. interior parts: Soffia Arnthorsdottir, personal comm.) Coarse scale orocesses: Sheep as a coarse-scale forager due to site fidelity (home range): For example, problem with re-growth on clear-cuttings if domestic animals turn the clear-cutting into inten- sively utilized grazing patches (faciiitating grass on expense of herbs and new birch tree stems sprouting from tree stumps); long- distance dispersal of moths (correlation length functions for bal- looning and female movements must be clarified for the model) Time: Fine-scale orocesses: Local insect outbreaks limited to old-age smaller stands and single trees Coarse-sc.ale nrocesses: Development of ungulate site fidelity to (and cultural transmission over generations) a local network of pre- ferred grazing patches. Direct anthropogenic and socioeconomic influence Space: Fine-sc.ale nrocesses: From mixed-age stands to more even-aged stand structure due to small clear-cuttings (loggings: firewood and forestry), Coarse scale orocesses: Distance to roads and villages (firewood and forestry: probability of a tree being logged at old age, and pro- bability of birch forest patch or stand mosaic); macroeconomic conditions for forestry; management plans with influence from offi- cial regulations and land use arrangements. Time: Fine scale orocesses: Episodic trampling effects on young birch shoots and seedlings at fine spatial scales Coarse scale orocesses: Shifting regulation policies for a region. "scale 2” special in this case? May the answer be revealed by exploring the relationship between spatio-temporal abun- dance pattern generated by the model under variable parameter settings for intrinsic growth rate, dispersal length and dispersal rate? These aspects among oth- ers need to be explored during the HIBECO modeling work, as an important validation of the birch-insect interactions. Table 1 gives some additional examples of scale-related chal- lenges for the model develop- ment. Scaling complexities emerge as a result of making the model sufficiently realistic (in accordance with definition above) by making it spatially explicit. Conclusion Spatially explicit models in gen- eral may often give unexpected results in comparison with their spatially implicit counterparts, and their greater level of realism make them a better starting point for validation against real systems like the mountain birch forest ecosystem. On the other hand a spatially explicit model puts huge demand on data, which also (at least for some parts) need to have specific spatially explicit details (e.g., insect outbreak series with spatial details, not just a local time series with a mean abundance variable). It is important to remember the old proverb for any modeler: "garbage in - garbage out". Thus, a model’s level of refinement and complexity must always be cali- brated and balanced in accor- dance with the quality and quan- tity of the available data. This is a prerequisite for model validation and production of scenario simu- lations with sufficient level of confidence. The actual HIBECO model development will be per- formed with this balance in mind. SKÓGRÆKTARRITIÐ 2001 l.tbl 149
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