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ment decision-making. But there are still two main limitations to most current DSTs: (1) firstly, most are not yet spatially explicit. i.e. they do not incorporate infor- mation at the landscape scale on the impacts of vegetation pattern and topography on ranging behaviour, partly because of a lack of data and partly because the software and computing power have only recently become sufficiently sophisticated; (2) sec- ondly, most of the available graz- ing models in the UK and else- where deal either only with open- ground vegetation or only with forest, because as yet we still have a poor understanding of for- est use when it represents only a small component of a whole complex of vegetation types. For this reason, a scoping studyis currently being undertaken to see where the main data shortages are, before we embark on new DST development for complex landscapes containing patches of forest within a ránge of other veg- etation types. This would aid managers of areas where trees form only a minor part of the landscape but where tree regen- eration is a major aim. Conclusion Grazing management for forest regeneration is a fundamentally important, yet highly complex, issue in countries such as Scotland, where natural forests have declined to such small remaining areas. Development of appropriate and sustainable management recommendations requires an integrated approach to a subject which encompasses many fields of expertise, from plant and animal physiology and ecology, through landscape sci- ence, to economics. it is only by integrating such approaches that we will be able to move towards the provision of sound prescrip- tions for a range of herbivore/for- est regeneration options within a changing economic and political climate. Acknowledgements I am grateful to Throstur Eysteinsson for inviting me to References Beaumont, D., Dugan, D., Evans, G., and Taylor, S. (1994) Deermanage- ment and tree regeneration in the RSPB reserve at Abernethy Forest. In: Aldhous, J.R. (Ed.), Our Pinewood Heritage. Forestry Commission, Edinburgh, pp. 186-195. Daneli, K., Niemela, P., Varvikko, T., and Vuorisala, T. (1991) Moose browsing on Scots pine along a gradient of plant productivity. Ecology, 72, 1624-1633. Danell, K„ Bergström, R„ and Edenius, L. (1994) Effects of large mammalian browsers on architecture, biomass, and nutrients of woody plants. J. Mammal., 75, 833-844. Edenius, L„ Danell, K. and Nyquist, H. (1995) Effects of simulated moose browsing on growth, mortal- ity, and fecundity in Scots pine. Can. J. For. Rcs„ 25, 529-535. Gilbert, D. this volume Gilbert, D„ Horsefield, D S-Thomp- son, D.B.A. (1997) The Ecology and Restoration of Montane and Sub- alpine Scrub Habitats in Scotland. Scottish Natural Heritage Review, 83, Scottish Natural Héritage, Battleby. Grace, J. & Easterbee, N. (1979) The natural shelter for red deer (Cervus elaphus) in a Scottish glen. Journal of Applied Ecology, 16, 37-48. Hester, A.|. (1995) Scrub in the Scottish Uplands. Scottisíi Natural Heritage RfWeu' No 24. Scottish Natural heritage, Battleby. Hester, A.|. & Miller, G.R. (1995) Scrub and woodland regeneration: prospects for the future. In Thompson, D.B.A., Hester, A.|. & Usher, M.B. (Eds) Heaths and Moorland: Cultural Landscapes. HMSO, Edinburgh. Pp 140-153. Hester, A.J., Mitchell, F.J.G. & Kirby, K.J. (1996) Effects of season and present this work at the FBT con- ference in lceland and for financ- ing my visit. Thanks also to Angela Sibbald for providing information and a Figure on the GPS project work (Fig 3). |ohn Milne provided helpful com- ments on the manuscript. intensity of sheep grazing on tree regeneration in a British upland woodland. For. Ecol. Manage., 88, 99-106. Mackenzie, N. (1999) The native woodland resource of Scotland. A review 1993-1998. Forestry Commis- sion Technica! Paper 30, Forestry Commission, Edinburgh. Millard, P„ Hester, A.|„ Wendler, R. & Baillie, G. (in press). Remobilization of nitrogen and the recovery of Betula pendula, Pinus sylvestris, and Sorbus aucuparia saplings aftersimulated browsing damage. Functional Ecology Mitcheil. B„ Staines, B.W. & Welch, D. (1977) Ecology of Red Deer. Institute of Terrestrial Ecology, Cambridge. MLURI (1998) MLURI Annual Report. MLURI, Aberdeen. Pastor, |. and Naiman, R. J. (1992) Selective foraging and ecosystem processes in boreal forests. Am. Nat„ 139, 691-705. Pastor,)., Moen, R.A., and Cohen, Y. (1997) Spatial heterogeneities, car- rying capacity, and feedbacks in animal-landscape interactions.). Mammal., 78, 1040-1052. Sibbald, A.M. (in press) Using GPS to track wild red deer stags. Deer. Staines, B.W. (1976) The use of natur- al shelter by red deer in north-east Scotland. lournal of Zoology, 180, 1-8. Stewart, F. E. (1996) The effects of red deer on the regeneration of upland birch woodland in the Scottish highlands. Unpublished PhD thesis, University of Aberdeen. Towers, this volume Van Hees, A. F. M„ Kuiters, A. T. and Slim, P. A. (1996) Growth and development of silver birch, pedunculate oak and beech as affected by deer browsing. For. Ecol. Manage., 88, 55-63. SKÓGRÆKTARRITIÐ 2001 l.tbl. 103
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